Kuja's Heritage
| Lion | Parents | Grand-Parents | Great Grand-Parents |
Great-Great Grand-Parents |
| Kuja | Kattalakis | ~Koyuki~ | Keyser Soze | Unknown |
| Unknown | ||||
| **QueenNanou** | Risal | |||
| z Bali | ||||
| Desoze | Keyser Soze | Unknown | ||
| Unknown | ||||
| Desenish | Shish Kebab | |||
| Desenoss | ||||
| Alenessa | π΅πππ | Boombah | Unknown | |
| [h] Aya (3268) | ||||
| Samekh | Boombah | |||
| Kill | ||||
| Du-ya | ~Koyuki~ | Keyser Soze | ||
| **QueenNanou** | ||||
| Rolling Muddy | Zeruaiah Nocturne | |||
| Klein | ||||
| This lion has 5 instances of possible inbreeding. Note: Inbreeding has no in-game effect. |
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